Evolution of ornamentation and pollination in Arecaceae with polymorphic species duplicated. A. Optimization of ornamentation type coded as 'Other-O' (white) and 'Psilate/Verrucate' (black). B. Optimization of pollination type coded as 'Other-P' (white) and 'Beetle' (black). The bicoloured branches indicate an equivocal inference of the ancestral character state. The transitions towards 'Beetle' pollination and 'Psilate/Verrucate' ornamentation are indicated by full crossbars and the reversals towards 'Other-P' pollination and 'Other-O' ornamentation are indicated by open crossbars (red and blue crossbars correspond respectively to the ACCTRAN and DELTRAN optimizations). Species names are coloured according to the subfamilies (Calamoideae in blue, Nypoideae in orange, Coryphoideae in green, Ceroxyloideae in pink and Arecoideae in red).