Table 1 Descriptive statistics for the three new species for 17 morphological characters

H. burgessi (19)

Average

18.5

6.4

5.8

6.5

1.0

1.9

1.9

3.8

2.4

9.2

8.8

5.9

7.7

4.4

5.0

5.0

5.1

St. Dev

1.3

0.4

0.4

0.5

0.1

0.1

0.1

0.5

0.2

0.5

0.5

0.3

0.5

0.4

0.2

0.4

0.4

Range

16.4-20.3

5.4-7.3

5.2-6.5

5.9-7.4

0.9-1.2

1.6-2.1

1.7-2.3

2.6-5

2.2-3

8.4-9.8

8-9.9

5.1-6.3

6.7-8.6

3.9-5.8

4.5-5.5

4.2-6.1

4.3-6.1

H. davenporti (10)

Average

18.9

6.5

5.4

6.4

1.0

1.9

1.9

3.8

2.4

9.1

9.1

5.8

7.4

4.4

5.4

5.4

5.0

St. Dev

1.0

0.3

0.3

0.4

0.1

0.1

0.1

0.2

0.2

0.4

0.4

0.4

0.7

0.2

0.3

0.3

0.4

Range

17.3-20.2

6-7.1

5-6.1

5.6-6.9

0.9-1.2

1.8-2.1

1.8-2

3.4-4

2.1-2.7

8.4-9.8

8.4-9.7

5-6.4

6.2-8.5

4.1-4.7

5.1-5.8

4.6-5.8

4.4-5.6

H. minutissimus (13)

Average

20.3

6.8

6.1

6.5

1.0

2.0

2.0

4.2

2.4

9.9

9.5

6.2

8.8

4.7

5.8

5.9

4.5

St. Dev

1.2

0.5

0.4

0.0

0.1

0.1

0.1

0.4

0.2

0.6

0.6

0.5

0.7

0.4

0.4

0.4

0.4

Range

18.8-22.7

6-7.5

5.5-6.8

6.5-6.5

0.9-1.1

1.7-2.1

1.7-2.3

3.4-4.7

1.9-2.8

8.7-11

8.6-10.7

5.5-7.3

7.8-10.3

4-5.5

5-6.8

5.2-6.9

3.7-5.1

H. spinigularis (9)

Average

19.6

6.9

6.1

6.9

1.1

1.9

2.0

3.9

2.5

9.9

8.8

6.2

8.2

4.5

5.3

5.1

4.0

St. Dev

1.0

0.4

0.4

0.4

0.1

0.1

0.2

0.2

0.3

0.3

0.6

0.4

0.7

0.2

0.4

0.4

0.3

Range

18.6-21.8

6.2-7.6

5.8-7.1

6.5-7.8

1-1.2

1.7-2.1

1.7-2.2

3.7-4.3

2-2.9

9.3-10.2

7.6-9.6

5.6-6.9

7.6-9.6

4.2-5

4.7-5.9

4.3-5.5

3.6-4.4

H. tanneri (2)

Average

22.2

6.9

6.1

6.9

1.2

2.0

2.1

4.1

2.7

11.1

8.9

6.0

9.5

4.7

6.3

NA

NA

St. Dev

0.4

0.9

0.6

0.5

0.1

0.2

0.3

0.8

0.4

0.7

1.6

1.1

1.1

0.4

0.7

NA

NA

Range

21.9-22.5

6.2-7.5

5.7-6.5

6.5-7.2

1.2-1.1

1.8-2.1

2.3-1.9

3.5-4.7

2.4-3

11.6-10.6

7.7-10

5.2-6.7

8.7-10.3

4.4-5

5.8-6.8

NA

NA

H. burgessi (28)

Average

24.3

8.5

7.2

8.3

1.3

2.4

2.3

4.8

3.1

12.1

11.2

7.6

10.3

5.4

7.1

St. Dev

1.2

0.5

0.6

0.7

0.1

0.1

0.3

0.3

0.3

0.8

0.9

0.4

0.7

0.3

0.4

Range

21.3-25.9

7.6-9.6

6.1-8.3

6.5-9.4

1-1.5

2.1-2.6

1.7-2.8

4-5.3

2.3-3.8

10.6-13.1

9.8-12.9

6.9-8.4

9-11.7

4.6-5.9

6-7.8

H. davenporti (2)

Average

27.0

9.2

6.9

8.4

1.3

2.3

2.6

5.2

2.8

12.9

13.0

8.6

10.7

6.3

7.4

St. Dev

0.1

0.0

0.0

0.3

0.1

0.1

0.1

0.2

0.1

0.4

0.6

0.1

0.3

0.7

0.2

Range

26.9-27

9.2-9.2

6.9-6.9

8.2-8.6

1.2-1.3

2.2-2.3

2.5-2.7

5-5.3

2.7-2.9

12.6-13.1

12.5-13.4

8.5-8.6

10.5-10.9

5.8-6.8

7.2-7.5

H. ukwiva (2)

Average

28.7

10.0

8.0

9.8

1.5

2.8

2.7

5.5

3.2

14.2

14.1

8.9

12.7

7.5

9.0

St. Dev

0.9

0.8

0.1

0.7

0.2

0.4

0.1

0.2

0.1

1.1

0.8

0.2

0.5

0.4

0.8

Range

28-29.3

9.4-10.5

7.9-8.1

9.3-10.3

1.3-1.6

2.5-3

2.6-2.8

5.3-5.6

3.1-3.3

13.4-15

13.5-14.6

8.7-9

12.3-13

7.2-7.7

8.4-9.6

H. spinigularis (3)

Average

24.5

8.7

7.3

8.8

1.3

2.3

2.6

4.9

3.1

12.9

11.8

8.2

11.1

5.6

7.0

St. Dev

1.1

0.6

0.5

0.6

0.0

0.1

0.1

0.1

0.3

0.3

0.6

0.5

0.8

0.4

0.4

Range

23.6-25.7

8.3-9.4

6.9-7.8

8.3-9.5

1.3-1.3

2.3-2.4

2.5-2.7

4.8-5

2.8-3.4

12.7-13.3

11.4-12.5

7.7-8.6

10.2-11.8

5.3-6

6.6-7.3

H. tanneri (2)

Average

26.7

9.2

7.6

8.7

1.5

2.4

2.7

5.2

2.9

15.0

11.2

7.7

10.7

6.1

7.4

St. Dev

4.3

1.5

1.1

0.7

0.3

0.5

0.2

0.5

0.4

5.7

3.3

2.3

3.0

1.7

1.5

Range

23.6-29.7

8.1-10.2

6.8-8.4

8.2-9.2

1.3-1.7

2-2.7

2.5-2.8

4.8-5.5

2.6-3.2

10.9-19

8.8-13.5

6.1-9.3

8.5-12.8

4.9-7.3

6.3-8.4